Subsection Seria Species Subspecies
Aristulatae Malz. Inaequaliglumis Malz. A.clauda Dur.  
A.pilosa M.B.
Stipitae Malz. A.longiglumis Dur.
A.ventricosa Balan. ventricosa (Balan) Malz.
bruhnsiana (Grun.) Malz.
Eubarbatae Malz. A.strigosa Schreb. strigosa (Schreb.) Thell.
hirtula (Lagas.) Malz.
barbata (Pott) Thell.
wiestii (Steud.) Thell.
vaviloviana Malz.
abyssinica (Hochst.) Thell.
Denticulatae Malz.   A.fatua L. septentrionalis Malz.
nodipilosa Malz.
meridionalis Malz.
macrantha (Hack.) Malz.
fatua (L.) Thell.
sativa (L.) Thell.
cultiformis Malz.
praegravis (Kraus.) Malz.
A.sterilis L. ludoviciana (Dur.) Gill.et Magn.
pseudo-sativa Thell.
trichophylla (C.K. et Hausskn.) Malz.
nodipubescens Malz.
macrocarpa (Monch.) Briq.
byzantina (C.K.) Thell.































1 Perennial plants A.macrostachya Bal.
  Annual plants 2
2 Lemma tips biaristulate 3
  Lemma tips bidentate 13
3 Glumes very unequal, lower glume one-half of upper one 4
  Glumes equal or nearly so 5
4 Disarticulate each floret at maturity A.clauda Dur.
  Disarticulation occurs at the lower floret only A.pilosa M.B.
5 Callus very long, awl shaped, 5-10 mm 6
  Callus short, about 2 mm 8
6 Disarticulate each floret at maturity A.longiglumis Dur.
  Disarticulation occurs at the lower floret only 7
7 Callus 5 mm long, glumes 25-30 mm long A.ventricosa Bal.
  Callus 10 mm long, glumes 40 mm long A.bruhnsiana Gruner.
8 Disarticulate each floret at maturity 9
  Panical non-shattering 13
9 Lemma tips biaristulate, glumes with 9-10 veins A.barbata Pott.
  Lemma tips biaristulate with 1-2 denticula or without, glumes with 7-9 veins 10
10 Lemma tips biaristulate with 1 denticulum, lemma tips longer than glumes; first floret scar narrow elliptic A.hirtula Lag.
  Lemma tips biaristulate with 2 denticula, lemma and glumes equal or nearly so; first floret scar oval 11
  Lemma tips biaristulate without denticula, first floret scar round 12
11 Lemma tips biaristulate 3-6 mm long A.wiestii Steud.
  Lemma tips biaristulate 1 mm long A.vaviloviana Mordv.
  Disarticulation occurs at the lower floret only A.atlantica Baum
12 Spikelets very small 12-15 mm A.prostrata Lad.
  Spikelets 20 mm long A.damascena Raj. et Baum
13 Lemma tips biaristulate with 1 denticulum, lemma and glumes equal and nearly so A.strigosa Schreb.
  Lemma tips biaristulate with 2 denticula,
lemma tips shorter than glumes
A. abyssinica Hochst.
14 All florets disarticulating at maturity 15
  Lowermost floret only disarticulating 16
  Panical non-shattering 19
15 Spikelets with 2-3 florets, glumes 15-20 mm long A.fatua L.
  Spikelets with 3-4 florets, glumes 25-30 mm long A.occidentalis Dur.
16 Spikelets very large with 3-5 florets, glumes 30-40 mm long 17
  Spikelets medium size with 2 rarely 3 florets, glumes 25-30mm long A.ludoviciana Dur.
  Spikelets small size with 2-3 florets, glumes 15-17 mm long A.canariensis Baum
  Spikelets very small size with 2 florets, glumes 13-15 mm long A.agadiriana Baum et Fedak
17 Lemma highly pubescent A.magna Murphy et Terr
  Lemma slightly-moderate pubescent A.sterilis L.
  Lemma pubescence is different 18
18 Point of awn insertion is about lower one-quarter, callus oval A.murphyi Lad.
  Point of awn insertion is between lower one-third and one-half, callus elliptical A.insularis Lad.
19 Fracture surface at the base of the primary floret is straighting A.sativa L.
  Fracture surface at the base of the primary floret is slanting A.byzantina C.Koch.
































Shape of panicle Hairiness of lemma Lemma colour A.strigosa
varieties
A.abyssinica
varieties
equilateral glabrous white albida Marq. chiovendae Mordv.
yellow   schimperi Korn.
grey typica Vav. hildebrantii Korn.
brown gilva Mordv. braunii Korn.
black melanocarpa Mordv.  
hairness of awn insertion white   typica Chiov.
yellow kewensis Vav.  
grey intermedia Marq.
black nigra Marq.
pubescent white alba Marq. solidiflora Thell.
grey-brown fusca Marq.  
flagged (brevis) glabrous white candida Mordv.
grey tephera Mordv.
brown hepatica Mordv.
black nigricans Mordv.
hairiness of awn insertion   semiglabra Malz.
pubescent trichophora Malz.
unilateral   secunda Mordv.
grains naked nudibrevis Vav.

Lemma colour Hairiness of lemma Awnedness Varieties
white Glabrous no awns albomutica Mordv.
Pubescent one awn alba Mordv.
cream Pubescent two awns cremea Mordv.
red-brown Glabrous no awns anolpa Mordv.
one awn rubra Mordv.
two awns culta Thell.
Pubescent one awn monathera Mordv.
two awns solida Haussk.
grey or black Glabrous one awn incana Mordv.
two awns nigra Mordv.
Pubescent one awn maroccana Mordv.
brown Glabrous one awn ursina Mordv.
Pubescent two awns cinnamomea Mordv.

Spikelets characters Lemma colour Awnedness Shape of panicle
Equilateral Unilateral
all florets thresh white no awns mutica Alef. obtusata Alef.
awns aristata Krause. tartarica Ard.
yellow no awns aurea Korn. flava Korn.
awns krausei Korn. ligulata Vav.
grey no awns grisea Korn. borealis Petrop.
awns cinerea Korn. armata Petrop.
brown no awns brunnea Korn. tristis Alef.
awns montana Alef. pugnax Alef.
the lowest floret threshes only white no awns volgensis Vav.  
awns kasanensis Vav.
yellow no awns segetalis Vav.
awns bashkirorum Vav.
brown awns armeniaca Mordv.
glumes up to 30 mm long white awns iranica Vav.
yellow awns persica Vav.
brown awns asiatica Vav.
lemma pubescent grey awns   homomala Mordv.
brown no awns setosa Korn.  
awns hausknechtii Vav.
grains naked white no awns inermis Korn.
awns chinensis Fisch.
white with no awns maculata Mordv.
dark spot awns mongolica Pissar.




























Subsection Species 2n Genom
Wild cultivated
floret disarticulation spikelet disarticulation
Aristulatae Malz. A.clauda Dur. A.pilosa M.B.   14 Cp
A.prostrata Ladiz.     Ap
A.damascena Raj.et Baum   Ad
A.longiglumis Dur.   Al
A.wiestii Steud. A.atlantica Baum   As
A.hirtula Lag.   A.strigosa Schreb.    
A.barbata Pott.   28 AB
A.vaviloviana Mordv. A.abyssinica Hoch.    
Denticulatae Malz.   A.ventricosa Bal.   14 Cv
A.bruhnsiana Grun.    
A.canariensis Baum   Ac
A.agadiriana Baum et Fed. 28 AB?
A.magna Mur.et Terr.   AC
A.murphyi Ladiz.    
A.insularis Ladiz.   CD?
A.fatua L. A.sterilis L. A.byzantina Koch 42 ACD
A.occidentalis Dur. A.ludoviciana Dur. A.sativa L.    



























Colour of lemma Hairiness of lemma Colour of hairiness Forms
white Glabrous   albida (Marq.)
Pubescent   albescens (Malz.)
grey (black) Glabrous   nigra (Marq.)
Pubescent   nigrescens (Malz.)
yellow Glabrous   flava (Korn.)
Pubescent   flavescens (Malz.)
brown Glabrous   braunii (Korn.)
Pubescent light genuina (Malz.)
dark atherantha (Malz.)